刚刚离开校园即将投身社会及工作的乡下姑娘安德丽娅·桑切丝(安妮·海瑟薇)幸运地得到了世上似乎所有女孩都梦寐以求的机会--在大城市纽约最出名时尚杂志《Runway》的主编米兰达·普雷斯丽(梅利尔·斯特里普)手下担任助理工作。如果助理工作做得好，安德丽娅将能留在这本整天与“Prada、Armani、Versace” 等世界著名服装设计师打交道的高贵杂志中担任羡煞旁人的编辑工作。当然，最重要的前提是她该如何能服侍好挑剔的主编米兰达，并获得她的最终肯定。身为著名时尚杂志的高级管理人，米兰达自身就有着高贵时尚、举止优雅的特点。这些无不令她手下的众人艳羡并敬畏不已。然而她这一看上去美好的一面却只对外人展示。在杂志社内部，米兰达绝对称得上是个不折不扣的“女魔头”。安德丽娅落在她手里堪称饱受折磨：首先，这位不时尚的姑娘每天早晨要应对米兰达的各种只说一遍的指令，并在每天第一时间给米兰达冲泡一杯香浓的咖啡，迟一秒钟都会被骂得狗血淋头；每天下班后还要接到工作狂米拉达打来的电话，应付她各种突如其来的工作和生活上的问题。当时，一心想得到米兰达肯定的安德丽娅虽有怨言但也任劳任怨。但是，在形式渐渐有所好转的情况下，她却突然发现，自己一直梦寐以求的工作原来并不如想象中的那么美好。倘若自己真的留在这家如地狱般的杂志社，有一天很可能会精神崩溃。这一刻，安德丽娅要作出如何的选择？
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