本片的剧情取自真实事件，故事来自于第二次世界大战期间的一个游击队传奇。克雷格所饰演的主角出生在波兰的一个犹太家庭里，身为长子的他很快成了家中的一把手。除他之外，家中还有两个弟弟，一家五口人其乐融融，生活倒也平静。但二战的炮火摧毁了一切！1939 年9 月1 日, 法西斯德**用”闪电战”的手段, 发动了侵略波兰的战争。仅10 多天时间, 就使波军全线溃败, 27 天后首都陷落, 国家灭亡。随着波兰政权的灭亡，纳粹对犹太人疯狂的屠戮紧跟其后。白色死亡的阴影笼罩大地，为了活命，身为长子的主角只身带领两个弟弟从纳粹德国占领的波兰逃往白俄罗斯。为了避开纳粹的追杀，最终遁入荒无人烟的大森林。
哪里有压迫哪里就有反抗，兄弟三人被当地的群众收留。在国仇家恨的推动下，他们义无反顾地加入前苏联反抗军，举起了抗击纳粹的大旗。俗话说得好，是金子总会发光的，在对抗纳粹出生入死的抗击过程中，主人公磨练出高超的作战技巧和沉稳冷静的性格，逐渐在他身边出现了一批跟随者。最终成长为一个抗击法西斯的游击队长，纳粹灭绝的是人性，对于出没于森林中的游击队，他们展开了疯狂的反扑。而主人公所率领的游击队则利用地形等因素，巧妙地与其周旋。丛林中的生活相当艰苦，危险时刻都会降临，但对自由的向往以及对纳粹的痛恨支持他们一步步走下去，甚至他们还在密林中建立起了一个他们自己的村庄！在二战期间，他们拯救了超过1200多条犹太人的生命，从而成就了一段传奇。
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